Simon and the Dai-Gurren brigade have toppled the Spiral King Lordgenome, and years later a far greater enemy, the Anti-Spiral, turns on a thriving humanity. Simon, Nia, and the team return to their giant mecha to meet it. This second of two compilation films retells the latter half of Tengen Toppa Gurren Lagann with heavily revised and newly animated footage, building to an expanded version of the series' climactic battle.