Tomoki Sakurai's quiet town life keeps fraying as Synapse, the floating world above the clouds, presses closer. The winged Angeloid Ikaros, bound to serve him, finds her connection to Tomoki running deeper than she understood, while Sohara, Mikako, and Sugata are drawn into the deepening conflict overhead. This feature film closes the Heaven's Lost Property story, following the original TV anime and its first film.