Eleven years after a cataclysm melted the Antarctic ice, raised the long-dead continent of Atlantis and woke the winged Fallen Angels, two-thirds of humanity is gone and the survivors are hunted for their life energy. Yurugi Kamishiro sits the pilot selection exam for the ultimate weapon Aquarion, a machine that forms when three excavated Vector Machines combine, passes it, and is still grinding through training when the regular pilots are wiped out and he is thrown into the fight. This manga tells its own story in the Genesis of Aquarion setting rather than retelling the TV series.