On Christmas Eve in Academy City, Touma Kamijou and the esper Mikoto Misaka decide to spend the holiday together, unaware that someone is quietly watching their every move. At the same time, the city's newly installed Board Chairman, Accelerator, sets in motion a plan that threatens to upend its entire power structure, stirring up enemies that had stayed hidden through the previous series. Genesis Testament continues the story begun in the New Testament novels, following Touma, Accelerator, and the Level 0 Shiage Hamazura as new fault lines open across the city.