Megumi Aino, a kind-hearted girl in the city of Pikarigaoka who is always looking to help others, is picked as a partner by Hime Shirayuki, a princess of the Blue Sky Kingdom. Hime already fights as Cure Princess, but she keeps running from battle against the Phantom Empire, the invaders led by Queen Mirage who conquered her kingdom and now attack Earth with Choiarks and Saiark monsters while Pretty Cure around the world fend them off. Blue, the spirit of Earth, gave Hime a Crystal of Love and told her to find someone to fight beside her, and Megumi becomes Cure Lovely.